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Yes, fatigue compromises form. But the good news is that as you get fitter you'll be able to sustain efficient form over longer periods of time. This simple fact is too often glossed over, as aspiring marathoners often want to see immediate training results. Based upon a runner's own athletic background and/or unique biomechanics, this may take time to develop.
Sunbeam spoke about loose toes, but loose shoulders, arms and hands are the keys to upper body running form. Our hands control the amount of tension in the upper body. So it's important to keep your hands in an unclenched fist, with fingers lightly touching palms. Should you feel your fists clenching or forearms tensing, drop your arms to your sides and shake them out for a few seconds to release the tension.
The same principal applies to shoulders. Carry them "low and loose", not "high and tight". Don't let them creep up toward your ears. Should this begin to happen, shake 'em out. I do this routinely throughout my long runs -- BEFORE tension can even begin to accrue.
Good luck in your marathon buildup.
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